Paleobiology , 24, Davidek, K. Palaeontographia Italica , 84, Download PDF extremely homemade, but better than nothing as original is difficult to find. Download PDF homemade. Journal of Paleontology , 71, Palaeontographica Canadiana , 14, Journal of Paleontology , 70, Geobios , 29, Journal of Paleontology , 69, Edgecombe, G.
Palaeontographica Abteilung A , , No PDF available sorry! Journal of Paleontology , 68, Journal of Vertebrate Paleontology , 14, Journal of Paleontology, 68, Canadian Journal of Earth Sciences , 30, Alcheringa , 16, Edgecombe and L. Smith Ebach, M.
The Devonian Trilobite Cordania from Australia. Wright NSW , Eldredge, N. Branisa Esker, G. Feist, R. Acta Palaeontologica Polonica, 47 2. Lerosey-Aubril The type species of Cyrtosymbole and the oldest Famennian cyrtosymboline trilobites. Klug, C. Schulz, and K. De Baets Red Devonian trilobites with green eyes from Morocco and the silicification of the trilobite exoskeleton.
Lerosey-Aubril, R. Feist Ontogeny of a new cyrtosymboline trilobite from the Famennian of Morocco. Osmolska, H. Acta Paleontologica Polonica, Vol. VIII, Number 4. Kin, and U. Radwanska Roy, S. Simoes, M. De Moraes Leme, and S. Soares XXI, Number 6. XIX, Number XIX, Number 1. XI, Number Trilobites of the Devonian Traverse Group of Michigan.
X, Number 6. Scripta Geologica, Wright, A. Haas A new Early Devonian spinose phacopid trilobite from Limekilns, New South Wales: morphology, affinities, taphonomy and palaeoenvironment. Records of the Australian Museum, 42 2. Amos, A. Campbell, and R. Goldring Australosutura Gen. Trilobita from the Carboniferous of Australia and Argentina. Goldring, R. Hahn, G.
Hahn Visean Trilobites from Holwell, Somerset. Brauckmann Lower Visean Trilobites from Feltrim, Ireland. Kandemir, R. Lerosy-Aubril Turkish Journal of Earth Sciences, Vol. Hamada Miller, J. Synonymy of the Carboniferous Trilobites Namuropyge and Coignouina. Ormiston, A. XIII, Number 3. Contributions to the Lower Carboniferous Cyrtosymbolinae Trilobita. XIII, Number 1. XII, Number 2.
Rak, S. Bulletin of Geosciences, 84 4. Grant, R. Advance Reports on the Permian Trilobites of Japan. Cordaniinae, nov. I - Outline of the Permian Trilobite Fauna. Angiolini Wass, R. Banks Some Permian Trilobites from Eastern Australia. Riek, E. Records of the Australian Museum, 26 I hope you don't mind me including them in the PDF library when you post them. That way they won't get buried underneath other posts as I continuously update.
You need to be a member in order to leave a comment. Sign up for a new account in our community. It's easy! Already have an account? Sign in here. Existing user? Sign in with Facebook. Recommended Posts. Fruitbat Posted July 19, Posted July 19, edited. Link to post Share on other sites. Fruitbat Posted July 24, Posted July 24, Updated July 24, Fruitbat Posted August 1, Posted August 1, Updated August 1, Fruitbat Posted August 6, Posted August 6, edited.
Updated and reorganized - August 7, Fruitbat Posted October 26, Posted October 26, Updated October 25, Fruitbat Posted November 19, Posted November 19, Updated November 19, Fruitbat Posted November 27, Frequency distributions of samples sizes for can Gondwana, North and South China, and Ordovician and Silurian collections designated in Table elsewhere.
The complete data set also includes 57 collections from detailed studies for which sample size is not known. TABLE 2. Significance levels of comparisons of frequen- cy distributions of E Sn for global data Figs. Significance levels for ularly for the Ordovician, we also included 57 nonrarefied data S are indicated only when conflict collections from detailed published studies in with the results for E Sn.
As a check on Series-level comparisons M-W K-S the potential bias introduced by differences in sample sizes, we repeated all analyses using a Shallow subtidal facies smaller subset of quantitative collections MO , UO 0. Quan- Ll 5 Lu S 0. In all analyses, we examined di- Deep subtidal facies versity patterns by comparing frequency dis- MO. Significance levels of comparisons of frequen- cy distributions of E Sn for series-level data split by cy distributions of E Sn for stage-level data of Laurentia province Figs.
Signifi- S tests. Significance levels for nonrarefied data S are cance levels for nonrarefied data S are indicated only indicated only where they conflict with the results for where they conflict with the results for E Sn. Late Rhuddanian—Aeronian Laurentia versus E. Avalonia 1 other E Sn 0. Telychian 0. M 1U Ordovician E Sn 0. Laurentia For most of the analyses, we divided the MO.
To evaluate the influence of the MO. Geographic occurrence of samples. A, Ordovician samples. B, Silurian samples. Shelf lithofacies used to classify collections according to habitat. Recent work has demonstrated that vician, and the new data show that this pat- there may be considerable geographic vari- tern continued into the Silurian. With one ex- ability in diversity patterns, both in the initial ception, modal species richness of both rare- recovery from mass extinction Jablonski fied and nonrarefied samples is in the 1—5 and in the background times between species class between the Middle Ordovician major events Miller Accordingly, we and Ludlow Fig.
Westrop and Ad- rain a found little variation in diversity and, to a lesser extent, East Avalonia in shal- between the Upper Cambrian and Upper Or- low subtidal and deep subtidal facies. However, in the expanded data set Table 2, Fig.
Series-level sample intervals used to examine temporal changes in trilobite species alpha diversity temporal changes in trilobite species alpha diversity. Frequency distributions of species richness and rarefied data [E Sn , at a standard sample size of 90 individuals] of all collections from Ordovician near- shore facies.
Di- versity is reduced in the Llandovery, but does not fall below the level of Middle Ordovician. Frequency distributions of species richness Ludlow diversity is indistinguishable from and rarefied data [E Sn , at a standard sample size of 90 Wenlock diversity, but is significantly lower individuals] of all collections from Ordovician and Si- lurian shallow subtidal facies.
Thus, results of statis- tical tests indicate progressive decline in al- pha diversity during the Silurian, so that the tinguishable from the Cambrian, even for the Llandovery and Ludlow are significantly dif- Wenlock and Ludlow Table 2.
Silurian data that can nificantly from the intervening Wenlock. The Westrop and Adrain a indicate that number of collections is small, but Silurian shallow-subtidal species richness was main- buildups are at least as diverse as those from tained at Late Cambrian levels during the the Middle Ordovician Fig. Most of the Si- Middle Ordovician but was significantly lurian collections are from the Wenlock, and greater than Cambrian levels in the Upper Or- these have species richnesses that are compa- dovician Table 2.
Frequency distributions of species richness, sample sizes, and rarefied data [E Sn , at a standard sample size of 90 individuals] of collections from Or- dovician and Silurian carbonate buildup facies. Frequency distributions of species richness, Cambrian and Middle Ordovician buildups sample size and rarefied data [E Sn , at a standard sam- from shelf-margin sites were more diverse ple size of 90 individuals] of collections from Ordovi- cian and Silurian deep subtidal facies.
Limit- ed data suggest that this pattern also holds for the Upper Ordovician Fig. The pattern of little a are indistinguishable from those of the net change between Upper Cambrian and Up- Upper Ordovician rarefied data only and per Ordovician deep subtidal collections Llandovery but are significantly lower in di- Westrop and Adrain a extends into the versity than the Middle Ordovician and Wen- Silurian Fig.
Diversity drops slightly be- lock Table 2. There are only a few collec- though it is significant in rarefied data only tions available from basinal facies, and none with a M-W test Table 2 ; with further decline include the relative abundance data required in the Llandovery, species richness is signifi- to perform rarefaction.
They display the same cantly lower than in both the Middle and Up- pattern as the nearshore facies: constant, low per Ordovician Table 2. By the Wenlock, di- species richness between the Middle Ordovi- versity had recovered and exceeded Middle cian and the Ludlow Fig. Ordovician levels Table 2; rarefied data only.
Mean values of species richness The number of collections from the Ludlow is for each habitat are presented in Figure 11, us- small but species richness is reduced dramat- ing S, rather than E Sn in order to maximize ically relative to the Wenlock.
Values are based on nonrarefied data. Cambrian data are from Westrop and Adrain a. Intercontinental Comparisons Shallow Subtidal Facies. Collections from the Middle and Upper Ordovician Fig. Frequency distribution of species richness those from Laurentia Table 3. In Laurentia, of collections from Ordovician and Silurian basinal fa- Middle and Upper Ordovician diversities are cies.
As also demonstrated Table 3 from the Cambrian data published using frequency distributions, there is no ev- by Westrop and Adrain a. There are no idence for changes in alpha diversity in the net changes between the Ordovician and Si- nearshore and basinal facies between the lurian of Laurentia: species richness in the Cambrian and the Silurian. In shallow and Llandovery is not significantly different from deep subtidal facies, maximum alpha diver- that in the Middle Ordovician Table 3 ; a sity is attained in the Upper Ordovician and small downward shift in modal species rich- Silurian, respectively, rather than the Cambri- ness in the Wenlock Fig.
In shallow subtidal facies, a decline in di- Table 3. There are few Ludlow collections, versity is evident in the Silurian following an but all fall in the upper half of the range for Ordovician peak, although it does not fall be- the Wenlock; combined data for the Wenlock low Cambrian levels. In the deep subtidal, Or- and Ludlow are indistinguishable from the dovician species richness exceeds Cambrian Middle Ordovician and Llandovery data Ta- levels, and there is a decline into the Llandov- ble 3.
There is Summary. Comparisons between various no significant difference in number of species paleocontinents are summarized in Figure 14, S between the Middle and Upper Ordovician in which species richness is expressed by combined data and the Llandovery Table 3. Other regions display levels of However, both Wenlock and Ludlow levels are Ordovician diversity that are comparable to below those for the Ordovician of Baltica Ta- those of Laurentia.
In Laurentia, the modest ble 3 and the Middle Ordovician of Laurentia post-Llandovery fall in shallow-subtidal spe- Table 3 ; Wenlock and Ludlow diversities of cies richness is not significant see above and Baltica are not significantly different Table 3.
Upper Cambrian of Baltica, but species rich- Equally sparse data from the Ludlow of East ness of Wenlock and Ludlow collections is not Avalonia show diversity minima in both shal- significantly different from that of collections low and deep subtidal settings. The decline in from the Upper Cambrian of Laurentia Table shallow-subtidal species richness in Baltica, 3. Deep Subtidal Facies. In the Ordovician, Response to the End-Ordovician Extinction peak diversity occurs in Laurentia, although At series-level resolution, there is only mut- all paleocontinents for which we have data ed response in the alpha diversity data to the show high species richness Fig.
Because end-Ordovician extinction: Llandovery shal- most of the deep subtidal data are from Lau- low-subtidal species richness is remarkably rentia, temporal changes for that continent are high in Laurentia Fig. To bring the pattern of diversity only. The number of collections However, the recovery of species richness ap- available for East Avalonia is small Fig. Frequency distributions of species richness upper set of histograms and rarefied data [E Sn , at a standard sample size of 90 individuals; lower set of histograms] of Ordovician and Silurian shallow subtidal col- lections from various paleocontinents.
For the Cambrian, data from Laurentia are from Westrop and Adrain a, and data from East Avalonia shallow subtidal are compiled from range charts in Rushton A only and Telychian Table 4 species richness small number of collections in the succeeding remaining below that of the Middle—Late Or- Telychian stage have the same mode as the dovician.
By the Wenlock, diversity had recov- Late Rhuddanian—Aeronian, and pooled data ered completely and actually exceeded Mid- for these two sample intervals are indistin- dle—Upper Ordovician levels Table 4; rarefied guishable from the Ordovician distribution data only. The apparent difference in the rate and the Wenlock Table 4. The recovery is, however, more pro- cies following the end-Ordovician extinction.
Frequency distributions of species richness upper set of histograms and rarefied data [E Sn , at a standard sample size of 90 individuals; lower set of histograms] of Ordovician and Silurian deep subtidal collections from various paleocontinents. Frequency distributions of species richness and rarefied data [E Sn , at a standard sample size of 90 individuals] of Laurentian collections from Ordovician and Silurian shallow subtidal facies left and deep subtidal facies right.
We significantly different between shallow and have commented earlier Adrain et al. Thus, the more Westrop and Adrain b upon the appar- extended period of recovery in deep subtidal ent decoupling of global clade and local eco- facies seems to be a by-product of the onshore- logical diversity, based on patterns evident in offshore diversity gradient in trilobite biofa- the Middle and Upper Ordovician.
The Silu- cies. Silurian diversity, even terns and global taxonomic diversity likely re- where there is evidence of a decline in species flects controls at hierarchical levels above richness in the Wenlock or Ludlow, did not those that regulate species richness in local drop below Upper Cambrian levels. Trilobite habitats. Determining the relative roles per Cambrian and the Upper Ordovician. We suggest that beta diversity is like- Ordovician and deep subtidal facies in the Si- ly to be the least important contributor.
Quan- lurian, rather than in the Cambrian. One of the titative studies of Upper Cambrian Ludvig- possibilities raised by Miller et al. The global sen biofacies reveal no marked changes survey presented herein, however, indicates in the degree of environmental partitioning. All regions for which data are available distribution patterns reflect a spectrum of bio- corroborate an increase in shallow-subtidal facies comparable to those that have been de- trilobite diversity from Middle to Upper Or- scribed from the Cambrian and Ordovician.
Certainly, there is no evidence for the kind of Silurian diversity trajectories do differ be- profound shift in patterns of habitat occupan- tween Laurentia and Baltica in shallow sub- cy that would be necessary to account fully for tidal facies, the only habitat type for which we the Ordovician drop in global diversity.
Differences in the pace of recovery down of biogeographic differences, especially from the end-Ordovician extinction between in shallow-water facies Owen Thus, a shallow and deep subtidal facies in Laurentia full understanding of diversity history of tri- appear to be the legacy of an offshore-directed lobites demands a hierarchical approach, and diversity gradient. The deep subtidal habitats available data suggest that changes in global that harbored more species before the extinc- taxonomic diversity of trilobites are not the re- tion took somewhat longer to regain preex- sult of ecological changes in local habitats.
Rather, movements of paleocontinents, along The results of this study provide further with Late Ordovician climatic deterioration, support for the suggestion Westrop et al. Article Google Scholar. Google Scholar. Denmark, 37 : —; Copenhagen. Balashova, E. Bondarev, V. Capera, J. Nord, 98 : 67—88; Lille. Dean, W. Part 1. Oslo, pp. Fortey, R. Remaining trilobites of the Valhallfonna Formation.
Skrifter, : 1—; Oslo. Howell, B. Hunt, A. Ingham, J. Lake, P.
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